Neurons and Action Potentials Study Pack

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Last updated May 27, 2026

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Neurons and Action Potentials Study Guide

Trace the full life cycle of an action potential — from resting membrane potential and the sodium-potassium pump to depolarization, repolarization, afterhyperpolarization, and saltatory conduction — so you can confidently explain every phase and the refractory period on your next exam.

Key Takeaways

  • Neurons maintain a resting membrane potential of approximately –70 mV, established by the sodium-potassium ATPase pump and the selective permeability of the membrane to potassium ions.
  • An action potential is an all-or-nothing electrical signal triggered only when membrane depolarization reaches the threshold of approximately –55 mV.
  • The rising phase of an action potential results from rapid opening of voltage-gated sodium channels, allowing Na⁺ to rush into the cell and drive the membrane potential toward +30 mV.
  • Repolarization occurs when voltage-gated sodium channels inactivate and voltage-gated potassium channels open, allowing K⁺ to exit the cell and restore the negative membrane potential.
  • A brief hyperpolarization (afterhyperpolarization) follows repolarization because potassium channels close slowly, temporarily making the membrane more negative than the resting potential.
  • Action potentials propagate along an axon without losing amplitude; in myelinated axons, this propagation is saltatory — the signal jumps between nodes of Ranvier, dramatically increasing conduction velocity.
  • The refractory period — divided into absolute and relative phases — prevents backward propagation of the signal and limits the maximum firing frequency of a neuron.

Neuron Structure and the Resting Membrane Potential

Before a neuron can transmit a signal, it must maintain a stable electrical state across its membrane — a condition called the resting membrane potential that depends on ion distribution and specialized transport proteins.

Functional Anatomy of a Neuron

  • The cell body (soma) contains the nucleus and metabolic machinery; dendrites receive incoming signals; the axon conducts output signals away from the soma.
  • Axons are often wrapped in myelin, a lipid-rich insulating sheath produced by Schwann cells in the peripheral nervous system and oligodendrocytes in the central nervous system.
  • Gaps in the myelin sheath called nodes of Ranvier are the only sites where voltage-gated ion channels are densely concentrated in myelinated axons.

Establishing the Resting Membrane Potential

  • The resting membrane potential of a typical neuron is approximately –70 mV, meaning the inside of the cell is negatively charged relative to the outside.
  • This charge difference arises because the membrane at rest is far more permeable to K⁺ than to Na⁺; potassium leaks outward through ungated leak channels, carrying positive charge with it.
  • The sodium-potassium ATPase pump actively transports 3 Na⁺ out of the cell and 2 K⁺ into the cell per cycle, consuming ATP to maintain steep concentration gradients for both ions.
  • Large, negatively charged proteins trapped inside the cell also contribute to the negative interior charge.

Threshold, Graded Potentials, and the All-or-Nothing Principle

Not every stimulus a neuron receives is strong enough to generate a full action potential; the membrane first integrates smaller, local voltage changes before deciding whether to fire.

Graded Potentials

  • A graded potential is a small, local change in membrane voltage caused by the opening of ligand-gated or mechanically gated ion channels at dendrites or the soma.
  • Unlike action potentials, graded potentials decay in amplitude with distance and can be either depolarizing (excitatory postsynaptic potentials) or hyperpolarizing (inhibitory postsynaptic potentials).
  • Multiple graded potentials can summate — either simultaneously (spatial summation) or in rapid succession (temporal summation) — at the axon hillock.

Threshold and the All-or-Nothing Rule

  • The axon hillock is the site of action potential initiation because it has the highest density of voltage-gated sodium channels in the neuron.
  • If summated depolarization raises the membrane potential to approximately –55 mV, this threshold is reached and an action potential fires.
  • The all-or-nothing principle states that once threshold is met, the action potential always reaches the same peak voltage regardless of how much the stimulus exceeded threshold; a stimulus below threshold produces no action potential at all.
  • Stimulus intensity is encoded not by the size of individual action potentials but by their frequency — stronger stimuli cause the neuron to fire more action potentials per second.

Phases of the Action Potential

An action potential unfolds as a precise, time-ordered sequence of ion channel openings and closings that produce a characteristic voltage spike lasting roughly 1–2 milliseconds.

Depolarization: The Rising Phase

  • At threshold, voltage-gated sodium channels open rapidly; the electrochemical gradient drives Na⁺ into the cell, making the interior progressively more positive.
  • This influx is self-reinforcing: depolarization opens more voltage-gated sodium channels, which causes more depolarization — a positive feedback loop that drives the membrane potential to approximately +30 mV.
  • At the peak, the interior of the cell is briefly positive relative to the exterior.

Repolarization: The Falling Phase

  • Voltage-gated sodium channels inactivate on their own shortly after opening, closing an inactivation gate and halting Na⁺ entry regardless of the membrane voltage.
  • Voltage-gated potassium channels, which open more slowly than sodium channels, are now fully open; K⁺ exits the cell down its electrochemical gradient, restoring the negative interior charge.
  • The membrane potential rapidly returns toward –70 mV as positive charge leaves the cell.

Afterhyperpolarization

  • Voltage-gated potassium channels do not close immediately when the resting potential is restored; they remain open briefly, allowing continued K⁺ efflux.
  • This drives the membrane potential slightly below the resting value to around –80 mV — a state called afterhyperpolarization or the undershoot.
  • The sodium-potassium ATPase pump works continuously to restore the original ion concentration gradients after repeated firing.

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